Chengjiang Biota
Fossils
 
Biota of the Maotianshan Shales, Chengjiang China


Also see:
Chengjiang Site Description
Chengjiang Fossils
Guanshan Fauna
Cambrian Explosion
Trilobite Relatives

 

The Chengjiang Maotianshan Shales known as ranks as one of the most important Lagerstätten fossil sites in the world. The Chengjiang site is located in the Yunnan Province of China in the villages of Ercaicun and Chengjiang near the city of Kunming. Some 50 meters of mudstone sediment are exposed, yielded many excellently preserved soft-bodied creatures of the Cambrian Explosion. Dated at 525 million years ago, it lies just above the Precambrian-Cambrian boundary at 544 million years ago.

Fossils of the Chengjiang China Maotianshan Shales
Chengjiang genera and species sorted by by kingdon or phylum:
Arthropoda - Arthropoda - Trilobitomorpha - Arthropoda - Class Trilobita - BrachiopodaChaetognatha - Cnidaria - Chordata - Ctenophora - Echinodermata - EnigmaticaHemichordata - Hyolitha - Lobopodia - Nematomorpha - Phoronida - Porifera - Protista - Priapulida - Uncertain - Vetulicolia


Phylum Arthropoda - 79 species
Acanthomeridion serratum (Hou et al., 1989)
Alalcomenaeus
Almenia spinosa (Hou & Bergstrom, 1997)
Apiocephalus elegans (Luo & Hu, 1999)
Branchiocaris? yunnanensis (Hou, 1987)
Canadaspis laevigata (Hou & Bergström, 1991)
Chengjiangocaris longiformis (Hou & Bergström, 1991)
Chuandianella ovata (Li, 1975)
Cindarella eucalla (Chen et al., 1996)
Clypecaris pteroidea (Hou, 1999)
Combinivalvula chengjiangensis (Hou, 1987)
Comptaluta inflata (Zhang, 1974)
Comptulata leshanensis (Lee, 1975)
Cyathocepalus bispinosus (Luo & Hu, 1999)
Dianchia mirabilis (Luo & Hu, 1997)
Diplopyge forcipatus (Luo & Hu, 1999)
Diplopyge minutus (Luo & Hu, 1999)
Dongshanocaris foliiformis (Hou & Bergström, 1998)
Ercaia minuscula (Chen et al.,2001)
Ercaicunia multinodosa
Forfexicaris valida (Hou, 1999)
Forticeps foliosa (Hou & Bergström, 1997)
Fuxianhuia protensa (Hou, 1987)
Glossocaris occulatus (Luo & Hu, 1999)
Guangweicaris spinatus (Guanshan Fauna)
Haikoucaris ercaiensis (Chen et al., 2004)
Isoxys auritus (Jiang, 1982)
Isoxys curvirostratus (Vannier & Chen, 2000)
Isoxys paradoxus (Hou, 1987)
Jianfengia multisegmentalis (Hou, 1987)
Jianshania furcatus (Luo & Hu, 1999)
Jiucunella paulula (Hou & Bergtrom, 1991)
Kuamaia lata (Hou, 1987)
Kuamaia muricata (Hou & Bergstrom, 1997)
Kuanyangia pustulosa (Lu, 1941)
Kunmingella angustacostata (Huo & Shu, 1983)
Kunmingella douvillei (Mansuy, 1912)
Kunmingella guanshanensis (huo & Shu, 1987)
Kunmingocaris bispinosus (Luo & Hu 1999)
Kunyangella cheni (Hou, 1965)
Leanchoilia asiatica (Luo & Hu, 1997)
Leanchoilia illecebrosa (Hou, 1987)
Liangshanella liangshenensis (Hou, 1956)
Mafangia subscalaria (Luo & Hu, 2002)
Mafangocaris multinodus (Luo & Hu, 2002)
Malongella bituerculata (Chang, 1974)
Occacaris oviformis Hou, 1999
Odaraia? eurypetala (Hou, 2004 to Pectocaris)
Ovalicephalis mirabilis (Luo & Hu, 2002)
Panlongia spinosa (Liu, 2006)
Panlongia tetranudosa (Liu, 2006)
Parakunmingella malongensis (Chang, 1974)
Parapaleomerus sinensis (Hou et al., 1999)
Pectocaris spatiosa (Hou, 1999)
Petalilium latus (Lupo & Hu, 1999)
Pisinnocaris subconigera (Hou & Bergström, 1998)
Primicaris larvaformis (Zhang & Hou, 2003)
Pseudoiulia cambriensis (Hou & Bergström, 1998)
Pterotum triacanthus (Luo & Hu, 2002)
Pygmaclypeatus daiensis (Zhang et al., 2000)
Rectifacies abnormalis (Hou et al., 1989)
Rhombicalvaria acanthi (Hou, 1987)
Saperion glumaceum (Hou et al., 1991)
Sidneyia sinica (Zang & Shu, 2002)
Sinoburius lunaris (Hou et al., 1991)
Skioldia aldna (Hou & Bergström, 1997)
Spinokunmingella typical (Huo & Shu, 1985)
Sunella grandis (Huo, 1965)
Squamacula clypeata (Hou & Bergström, 1997)
Syrrhaptis intestinalis (Luo & Hu, 1999)
Urokodia aequalis (Hou et al., 1989)
Tanglangia caudata (Luo & Hu, 1999)
Trigoides aclis (luo & Hu, 1999)
Tsunydiscus aclis (Zhou, 1975)
Tsunyiella daindongensis (Tong, 1985)
Tuzoia sinensis - a Phyllocarid (Pan, 1957)
Waptia ovata (Lee, 1975)
Wutingella binodosa (Zhang, 1974)
Xandarella spectaculum (Hou et al.,1991)
Yiliangocaris ellipticus (Luo & Hu, 1999)
Yunnanocaris megista (Hou, 1999)

Phylum Arthropoda - Trilobitamorpha to Arachnomorpha
- 2 species of Naraoiids (Cotton & Braddy, 2004)
Naraoia Misszhouia longicaudata (Zhang & Hou, 1985)
Naraoia spinosa (Zhang & Hou, 1985)

Phylum Arthropoda - Class Trilobita - 5 species
Eoredlichia intermedia; Family Redlichiidae (Lu, 1940)
Kuanyangia pustulosa; Family Redlichiidae (Lu, 1941)
Wutingaspis tingi; Family Redlichiidae (Kobayashi, 1944)
Yunnanocephalus yunnanensis; Family Yunnanocephalidae (Mansuy, 1912)
Palaeolenus lantenoisi Family Palaeolenidae (Mansuy, 1912)

Phylum Brachiopoda - 5 species
Diandongia pista (Rong, 1974)
Heliomedusa orienta (Sun & Hou, 1987)
Lingulella chengjiangensis (Jin et al., 1993)
Lingulellotreta malongensis (Rong, 1974)
Longtancunella chengjiangensis (Hou et al., 1999

Phylum Chaetognatha - 1 species
Eognathacantha ercainella (Chen & Huang, 2002)

Phylum Cnidaria - 2 species
Priscapennamarina angusta (? Zhang & Babcock, 2001)
Xianguangia sinica (Chen & Erdtmann, 1991)

Phylum Chordata - 8 species
Cathaymyrus diadexus (Shu, et al., 1996)
Cathaymyrus haikouensis (? Luo & Hu, 2001)
Haikouella jianshanensis ( Shu, et al., 2003)
Haikouella lanceolata (Chen et al., 1999)
Haikouichthys ercaicunensis (Leo, et al. 1999)
Myllokunmingia fengjiaoa (Shu et al., 1999)
Shancouclava anningense - aTunicate- (Chen et al., 2003)
Zhongxiniscus intermedius (? Luo & Hu, 2001)

Phylum Ctenophora - 3 species
Maotianoascus octonarius Chen & Zhou, 1997
Sinoascus paillatus (Chen & Zhou, 1997)
Stromatoveris psygmoglena (?) (Shu, 2006)

Phylum Echinodermata - 1 species
Cotyledion tylodes

Enigmatic animals - 24 animals
Allonnia phrixothrix (Bengtson & Hou, 2001)
Amiskwia sinica (Luo & Hu, 2002)
Anthrotum robustus (Luo & Hu, 1999)
Batofasciculus ramificans (Hou et al., 1999)
Cambrotentacus sanwuia (Zhang & Shu, 2001)
Conicula straita (Luo & Hu, 1999)
Dinomischus venustus (Chen et al., 1989)
Discoides abnormalis (Luo & Hu, 1999)
Eldonia eumorpha (Sun & Hou, 1987)
Facivermis yunnanicus (Hou & Chen, 1989) Putative Lobopod (Liu, 2006)
Hippotrum spinatus (Luo & Hu,1999)
Jiucunia petalina (Hou et al., 1999)
Maanshania crusticeps (Hou et al., 1999)
Macrocephalus elongates (Luo & Hu, 1999)
Parvulonoda dubia (Rigby & Hou, 1995)
Phacatrum tubifer (Luo & Hu, 1999)
Phasangula striata (Luo & Hu, 1999)
Phlogites brevis (Luo & Hu, 1999)
Phlogites longus (Luo & Hu, 1999)
Priscapennamarina angusta (Zhang & Babcock, 2001)
Pristitoites bifarius (Luo & Hu, 1999)
Rhipitrus calvifer (Luo & Hu, 1999)
Rotadiscus grandis (Sun & Hou, 1987)
Yunnanozoon lividum (Hou et al., 1991) - putative earliest known hemichordate

Phylum Hemichordata - 1 species
Galeaplumosus abilus

Phylum Hyolitha - 8 species
Ambrolinevitus maximus (Jiang, 1982)
Ambrolinevitus platypluteus (Jaing, 1982)
Ambrolinevitus ventricosus (Qian, 1978)
Burithes yunnanensis (Hou et al., 1999)
Glossolithes magnus (Luo & Hu, 1999)
Linevitus billingsi (Qian, 1989)
Linevitus flabellaris (Qian, 1978)
Linevitus opimus (Yu, 1974)

Phylum Lobopodia - 6 species
Cardiodictyon catenulum (Hou et al., 1991)
Hallucigenia fortis (Hou & Bergström, 1995)
Luolishania longicruris (Hou & Chen, 1989)
Microdictyon sinicum (Chen et al., 1989)
Miraluolishania haikouensis
Onychodictyon ferox (Hou et al., 1991)
Paucipodia inermis (Chen et al., 1995)

Phylum Nematomorpha - 3 species
Cricocosmia jinningensis (Hou & Sun, 1988)
Maotianshania cylindrica (Sun & Hou, 1987)
Palaeoscolex sinensis (Hou & Sun, 1988)

Phylum Phoronida - 1 species
Iotuba chengjiangensis (Chen & Zhou, 1997)

Phylum Porifera - 15 species
Allantospongia mica (Rigby & Hou, 1995)
Choia xiaolantianensis (Hou et al., 1999)
Choiaella radiata (Rigby & Hou, 1995)
Hazelia sp.
Leptomitella confusa (Chen et al., 1989)
Leptomitella conica (Chen et al., 1989)
Leptomitella metta (Rigby, 1983)
Leptomitus teretiusculus (Chen et al., 1989)
Paraleptomitella dictyodroma (Chen et al., 1989)
Paraleptomitella globula (Chen et al., 1989)
Quadrolaminiella crassa (Chen et al., 1990)
Quadrolaminiella diagonalis (Chen et al., 1990)
Saetaspongia densa (Mehl & Reitner, 1993)
Sinfoflabrum antiquum (? Zhang & Babcock, 2001)
Triticispongia diagonata (Mehl & Reitner, 1993)

Kingdom Protista (algae) - 4 species
Fuxianospira gyrata (Chen & Zhou, 1997)
Megaspirellus houi (Chen & Erdtmann, 1991)
Sinocylindra yunnanensis (Chen & Erdtmann, 1991)
Yuknessia sp. (Chen & Erdtmann, 1991)

Phylum Priapulida - 16 species
Acosmia maotiania (Chen & Zhou, 1997)
Archotuba conoidalis Hou et al., 1999
Corynetis brevis (Luo & Hu, 1999)
Gantoucunia aspera (Luo & Hu, 1999)
Lagenula triolata (Luo & Hu, 1999)
Oligonodus specialis (Luo & Hu, 1999)
Paraselkirkia jinningensis (Hou et al., 1999)
Palaeopriapulites parvus (Hou et al., 1999)
Protopriapulites haikouensis (Hou et al., 1999)
Sandaokania latinodosa (Luo & Hu, 1999)
Selkirkia elongata (Luo & Hu, 1999)
Selkirkia sinica (Luo & Hu, 1999)
Sicyophorus rarus (Luo & Hu, 1999)
Xiaoheiqingella peculiaris (Luo & Hu, 2002)
Xishania longgiusula (Lup & Hu, 2002)
Yunnanopriapulus halteroformis (Huang & Chen, 2004)

Phylum Uncertain - 4 species
Amplectobelua symbrachiata (Hou et al., 1995)
Anomalocaris saron (Hou et al., 1995)
Cucumericrus decoratus (Hou et al., 1995)
Parapeytoia yunnanensis (Hou et al., 1995)

Phylum (?) Vetulicolia - 7 species
Banffia confusa (Chen & Zhou, 1997)
Didazoon hoae (Shu & Han, 2001)
Vetulicola cuneata (Hou, 1987)
Vetulicola gantoucunensis (Luo, et al, 2005)
Vetulicola rectangulata (? Luo & Hou, 1999)
Xidazoon stephanus (Shu, et al., 1999)
Yuyuanozoon magnificissimi (Chen, et al.,2003)

References

Artribution: Phyla Genera and Species of the Maotianshan Shales Chengjiang Biota